Olethreutinae

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Tortricinae

H

Epiblema males compared (provisional)

uddmanniana

scutulana/cirsiana

costipunctana

N broad, same width as S
C = strongly tapering upwards.
H = strong. P = lacking

N broad, similar width as S
C = parallel, oblong. H = small
P = moderate

N broad, similar width as S
C = narrow, elliptical. H = tiny, but large semi-circular bristly pad (pulvinus). P = large

C

S

N

P

pulvinus

foenella

roborana

incarnatana

N narrow. S broad and angular (A)
C = slightly tapering upwards. H = moderate. P = large, taller than H.

N long,broad, same width as S
C = slightly tapering upwards, narrow. Margin (M) scalloped.

N short,broad, same width as S
C = elliptical, broad. Margin (M) scalloped.

A

C

N

S

P

M

S = sacculus; C = cucullus; N = neck; H = horn;
P = prominence; A = angle; M = margin; L = lobe

Epiblema or Eucosma?

cynosbatella

N squat, wider than S
C = broad, ventral lobe (L) lacking. H = small, triangular.

C

N

S

H

L

trimaculana*

rosaecolana*

N very narrow. S = fairly broad
C = slightly tapering upwards. Lobe (L) pointed. H = moderate. P = strongly curved. Socius (sc) long

N very narrow. S = fairly narrow
C = parallel. Lobe (L) rounded. H = moderate. P = weakly curved.
Socius (sc) short

P

sc

sc

H

sticticana

N narrow. S = fairly broad, angular (A). C = parallel. H =missing, no apparent pulvinus. P = strongly arched.

A

C

N

S

* trimaculana/rosaecolana
full male genitalia compared

cynosbatella I uddmanniana I trimaculana I rosaecolana I roborana I incarnatana I foenella I sticticana I costipunctana

Epiblema or Eucosma?